Shawn Fraver – Selected Publications
Link to Google Scholar Page: https://scholar.google.com/citations?hl=en&user=snx7NFAAAAAJ
Note: Student or Post-doc, if first author
2024 (to date)
Fraver, S., C. Bosely-Smith, C.E. Sierup, C.H. Guiterman, T. Schmeelk, A. Teets, R. VanKampen,
L.S. Kenefic. Dendrochronological reconstruction of arborvitae leafminer (Argyresthia spp.)
outbreaks on northern white-cedar (Thuja occidentalis) in Maine, USA. In press, Canadian
Journal of Forest Research. https://doi.org/10.1139/cjfr-2023-0193
Jati, A., B.W. Broto, G. Franzoi Dri, K. Latifiana, S. Fraver, I. Sri Rejeki, Bustang, A. Mortelitti.
2024. Conserving large mammals on small islands: a case study on one of the world’s most
understudied pigs, the Togean Islands babirusa. Biodiversity and Conservation.
https://doi.org/10.1007/s10531-024-02800-5
Ray, D.G., R.S. Seymour, S. Fraver, L. Kenefic, P. Berrill, N. Rogers, A.R. Weiskittel. 2024.
Quantifying minimum site occupancy requirements of common forest tree species in
northern New England, USA: Implications for stocking assessment. Forest Ecology and
Management 566. 122057
Zanne, A., S. Allison, W. Baptiste, J.H.C Cornelisson, W. Cornwell, P. Eggleton, S. Fraver, M. Ooi, J. Powell, C.W. Woodall. 2024. The role of dead wood in the carbon cycle: implications for models, forest management, and future climates. Annual Review of Ecology, Evolution, and Systematics, 55.
2023
Read, Z., S. Fraver, A.W. D’Amato, D. Evans, K. Evans, D. Lutz, C.W. Woodall. 2023. CO 2 flux from Acer saccharum logs: Sources of variation and the influence of silvicultural treatments.
Canadian Journal of Forest Resources. 53(8): 654–662. DOI: 10.1139/cjfr-2022-0291.
Bosely-Smith, C., S. Fraver, J. Wason, N. Rogers, A.W. D’Amato. 2023. The natural ‘exclosure
effect’ and tree regeneration following post-windstorm salvage logging. Journal of Applied
Ecology. DOI: 10.1111/1365-2664.14560.
Ray, D.G., R.S. Seymour, S. Fraver, J.-P. Berrill, L. Kenefic, N. Rogers, A.R. Weiskittel. 2023.
Relative density as a unifying principle for density management diagrams and stocking
guides. Journal of Forestry 121: 443–456. DOI: https://doi.org/10.1093/jofore/fvad029
Forrester, J.A., S. Fraver, D.J. Mladenoff, S.T. Gower, A.W. D’Amato, D.L. Lindner. 2023.
Experimental evidence that forest structure controls detrital decomposition. Ecosystems.
https://doi.org/10.1007/s10021-023-00841-5.
French, K., M.A. Vadeboncoeur, H. Asbjornsen, S. Fraver, L.S. Kenefic, D.B. Moore, J.W. Wason. 2023. Temporary thinning shock in previously shaded red spruce. Canadian Journal of Forest Resources 53(7): 491–502. DOI: 10.1139/cjfr-2022-0227.
Perreault, L., J.A. Forrester, D.L. Lindner, M.A. Jusino, S. Fraver, M.T. Banik, D.J. Mladenoff.
2023. Linking wood-decay fungal communities to decay rates: Using a long-term
experimental manipulation of deadwood and canopy gaps. Fungal Ecology 62: 101220.
doi.org/10.1016/j.funeco.2022.101220.
Käber, Y., … S. Fraver, et al. 2023. Sheltered or suppressed? Tree regeneration in unmanaged
European forests. Journal of Ecology 111(10): 2281-2295. DOI: 10.1111/1365-2745.14181
2022
Fraver, S., A.W. D’Amato, M. Reinikainen, K.G. Gill, B.J. Palik. 2022. Dynamics and structure of
old-growth Fraxinus nigra stands in northern Minnesota. Canadian Journal of Forest
Research 52 (6): 910–919. https://doi.org/10.1139/cjfr-2021-0340
Read, Z., S. Fraver, J.A. Forrester, J. Wason, C.W. Woodall. 2022. Temporal trends in CO 2 flux
from Picea rubens stumps: A chronosequence approach. Forest Ecology and Management
524: 120528. https://doi.org/10.1016/j.foreco.2022.120528
Green, M., S. Fraver, D. Lutz, C. Woodall, A.W. D’Amato, D. Evans. 2022. Does deadwood
moisture vary jointly with surface soil water content? Soil Science Society of America
Journal. DOI: 10.1002/saj2.20413
Dagley, C.M., J.-P. Berrill, S. Fraver. 2022. Forest restoration mitigates drought vulnerability of
coast Douglas-fir in a Mediterranean climate. Canadian Journal of Forest Resources 53(3):
210-216. https://doi.org/10.1139/cjfr-2022-0119
Teets, A., D.J.P. Moore, M.R. Alexander, P.D. Blanken, G. Bohrer, S.P. Burns, M.S. Carbone, M.J.
Ducey, S. Fraver, C.M. Gough, D.Y. Hollinger, W. Munger, K.A. Novick, S.V. Ollinger, A.P.
Ouimette, N. Pederson, B. Seyednasrollah, C.S. Vogel, A.D. Richardson. 2022. Coupling of
tree growth and photosynthetic carbon uptake in six North American forests. Journal of
Geophysical Research: Biogeosciences 127, e2021JG006690. https://doi.org/10.1029/2021JG006690.
2021
Allogio, J., S. Fraver, L. Kenefic, J. Wason, P. Berrill. 2021. Regeneration and recruitment in
lowland northern white-cedar (Thuja occidentalis) forests. Forest Ecology and Management
499: 119639
Woodall, C.W., S. Fraver, S.N. Oswalt, S.A. Goeking, G.M. Domke, M.B. Russell. 2021. Decadal
dead wood biomass dynamics of coterminous US forests. Environmental Research Letters
16, 104034.
Hollinger, D.Y., E. A. Davidson, S. Fraver, H. Hughes, J.T. Lee, A.D. Richardson, K. Savage, D. Sihi,
and A. Teets. 2021. Multi-decadal carbon cycle measurements indicate resistance to external drivers of change at the Howland Forest AmeriFlux site. Journal of Geophysical
Research – Biogeosciences. 126, e2021JG006276.
Gleason, K.E, J.B. Bradford, A.W. D’Amato, S. Fraver, B. Palik, M.A. Battaglia. 2021. Forest
density intensifies the importance of snowpack to growth in water‐limited pine forests.
Ecological Applications 31(1): e02211.
Wei, X., D.J. Hayes, I. Fernandez, S. Fraver, J. Zhao, and A. Weiskittel. 2021. Climate and
atmospheric deposition drive the inter-annual variability and long-term trend of dissolved
organic carbon flux in the conterminous United States. Science of the Total Environment
771: 145448.
Wei, X., D.J. Hayes, I. Fernandez, J. Zhao, S. Fraver, C. Chan. 2021, J. Diao. 2021. Identifying key
environmental factors explaining temporal patterns of DOC export from watersheds in the
conterminous United States. Journal of Geophysical Research – Biogeosciences 126:
e2020JG005813.
Ayrey, E., D.J. Hayes, J.B. Kilbride, S. Fraver, J.A. Kershaw Jr., B.D. Cook, A.R. Weiskittel. 2021.
Synthesizing disparate LiDAR and satellite datasets through deep learning to generate wall-
to-wall regional forest inventories for the complex, mixed-species forests of the eastern
United States. Remote Sensing 13, 5113.
2020
Fraver, S., L. Kenefic, A. Cutko, A.S. White. 2020. Natural disturbance and stand structure in old-
growth Thuja occidentalis forests of northern Maine. Forest Ecology and Management 456:
117680.
Costanza, K.K.L., W.H. Livingston, S. Fraver, I.A. Munck. 2020. Dendrochronological analyses
and whole tree dissections reveal Caliciopsis canker (Caliciopsis pinea) damage associated
with declining growth and climatic stressors of eastern white pine (Pinus strobus). Forests.
11: 347.
Woodall, C., D. Evans, S. Fraver, M. Green, D. Lutz, A.W. D’Amato. 2020. Real-time monitoring
of dead wood moisture and temperature in forests: Lessons learned from an intensive case
study. Canadian Journal of Forest Research 50: 1244–1252.
Andrews, C.A., A.W. D’Amato, S. Fraver, B. Palik, M.A. Battaglia, J.B. Bradford. 2020. Stand
density moderates growth declines during hot-droughts in dry forests. Journal of Applied
Ecology 57(6): 1089-1102.
2019
Fien, E.K.P., S. Fraver, A. Teets, A.R. Weiskittel, D.Y. Hollinger. 2019. Drivers of individual tree
growth and mortality in an uneven-aged, mixed-species conifer forest. Forest Ecology and
Management 449: 117446.
Puhlick, J.J., S. Fraver, I.J. Fernandez, A. Teets, L. Kenefic, A.R. Weiskittel. 2019. Site quality,
disturbance, and vegetation effects on carbon storage and accumulation in old, mixed-
species stands in central Maine, USA. Natural Areas Journal 39(4): 429-441.
Woodall, C.W., V.J. Monleon, S. Fraver, H. Hatfield, J. Campbell, G.M. Domke, M. Russell. 2019.
Data descriptor: The downed woody materials inventory of United States Forests. Nature
Scientific Data 6: 180303. DOI: 10.1038/sdata.2018.303.
Dodds, K., M.F. DiGirolomo, S. Fraver. 2019. Response of bark beetles and woodborers to
tornado damage and subsequent salvage logging in northern coniferous forests of Maine,
USA. Forest Ecology and Management 450: 117489.
Campbell, J.L., M.B. Green, R.D. Yanai, C.W. Woodall, S. Fraver, C.R. See, G.M. Domke, M.E.
Harmon. 2019. Estimating uncertainty in the volume and carbon storage of downed coarse
woody debris. Ecological Applications 29(2):e01844.
Ayrey E., D.J. Hayes, S. Fraver, J.A. Kershaw, A.R. Weiskittel. Ecologically-based metrics for
assessing structure in developing area-based, enhanced forest inventories from LiDAR.
Canadian Journal of Remote Sensing 45(1): 88-112.
Mortelliti, A., I. Grentzmann, S. Fraver, S. Calkins, N. Fisichelli, A.M. Brehm. 2019. Predicting the
climate-induced expansion of woody plant species by identifying critical biological controls.
Oikos 128: 1726-1738.
Ayrey, E., D.J. Hayes, J.B. Kilbride, S. Fraver, J.A. Kershaw Jr., B.D. Cook, A.R. Weiskittel. 2019.
Synthesizing disparate LiDAR and satellite datasets through deep learning to generate wall-
to-wall regional forest inventories. BioRxiv: 580514.
2018
Fraver, S., M.J. Ducey, C.W. Woodall, A.W. D’Amato, A.M. Milo, B.J. Palik. 2018. Influence of
transect length and downed woody debris abundance on precision of the line-intersect
sampling method. Forest Ecosystems 5: 39. https://doi.org/10.1186/s40663-018-0156-9.
Fraver, S., M. Tajvidi, D.L. Lindner, A.W. D’Amato, J.A. Forrester, A.M. Milo. 2018. Woody
material structural degradation through decomposition on the forest floor. Canadian
Journal of Forest Research 48: 111-115.
Teets, A., S. Fraver, D. Hollinger, A. Weiskittel, R.S. Seymour, A. Richardson. 2018. Linking
annual tree growth with eddy-flux measures of net ecosystem productivity across twenty
years of observation in a mixed conifer forest. Agricultural and Forest Meteorology 249:
479-487.
Teets, A., S. Fraver, A. Weiskittel, D. Hollinger. 2018. Quantifying climate – growth relationships
at the stand-level in a mature mixed-species conifer forest. Global Change Biology 24: 3587-
3602.
Wesely, N., S. Fraver, L.S. Kenefic, J.-C. Ruel, A. Weiskittel, M. Thompson, A. White. 2018.
Structural attributes of old-growth and partially harvested northern white-cedar stands in
Northeastern North America. Forests 9, 376: 1-15.
Ducey, M.J., S. Fraver. 2018. The conic-paraboloid formulae for coarse woody material volume
and taper, and their approximation. Canadian Journal of Forest Research 48: 966-975.
Schurman, J.S., V. Trotsiuk, V. Čada, S. Fraver, P. Janda, D. Kulakowski, T. Nagel, R. Seidl, M.
Svoboda. 2018. Large-scale disturbance legacies and the climate sensitivity of primary Picea
abies forests. Global Change Biology 24(5): 2169-2181.
D’Orangeville, L., J. Maxwell, D. Kneeshaw, N. Pederson, L. Duchesne, T. Logan, D. Houle, D.
Arseneault, D.A. Bishop, D. Druckenbrod, S. Fraver, F. Girard, J. Halman, C. Hansen, J.L. Hart,
H. Hartmann, M. Kaye, D. Leblanc, S. Manzoni, S. Rayback, C. Rollinson, R.P. Phillips. 2018.
Timing and local water balance regulate drought impacts on the temperate forest carbon
sink. Global Change Biology 24(6): 2339-2351.
Sommerfeld, A., C. Senf, B. Buma, A.W. D’Amato, T. Després, I. Díaz-Hormazabal, S. Fraver, L.E.
Frelich, Á.G. Gutiérrez, S.J. Hart, B.J. Harvey, H.S. He, T. Hlásny, A. Holz, T. Kitzberger, D.
Kulakowski, D. Lindenmayer, A.S. Mori, J. Müller, J. Paritsis, G. Perry, S. Stephens, M.
Svoboda, M.G. Turner, T.T. Veblen, R. Seidl. 2018. Recent patterns of natural disturbance in
the temperate forest biome. Nature Communications 9: 4355.
Looney, C.E., A.W. D’Amato, S. Fraver, J.B. Bradford, D. Kastendick, B.J. Palik. 2018. Size-growth
relationship, tree spatial patterns, and tree-tree competition influence tree growth and
stand complexity in a 160-year red pine chronosequence. Forest Ecology and Management
424: 85-94.
Marra, R., N. Brazee, S. Fraver. 2018. Estimating carbon loss due to internal decay in living trees
using tomography: implications for forest carbon budgets. Environmental Research Letters
13: 105004.
Trotsiuk, V., D. Druckenbrod, D. Martin-Benito, D. Orwig, D. Bishop, A. Barker-Plotkin, S. Fraver,
N. Pederson. 2018. Testing the efficacy of tree-ring methods for detecting past
disturbances. Forest Ecology and Management 425: 59-67.
Sass, E.M., A.W. D’Amato, D.R. Foster, A. Barker Plotkin, S. Fraver, P.K. Schoonmaker, D.A.
Orwig. 2018. Long-term influence of disturbance-generated microsites on forest structural
and compositional development. Canadian Journal of Forest Research 48: 966-975.
Leverkus, A.B., J.M.R. Benayas, J. Castro, D. Boucher, S. Brewer, B.M. Collins, D. Donato, S.
Fraver, B.E. Kishchuk, E.-J. Lee, D. Lindenmayer, E. Lingua, E. Macdonald, R. Marzano, C.C.
Rhoades, S. Thorn, A. Royo, J.W. Wagenbrenner, K. Waldron, T. Wohlgemuth, L. Gustafsson.
2018. Salvage logging effects on regulating and supporting ecosystem services. Canadian
Journal of Forest Research 48: 983-1000.
D’Amato, A.W., P. Raymond, S. Fraver. 2018. Old-growth disturbance dynamics and associated
ecological silviculture for forests in northeastern North America. IN: Eastern Old-Growth
Forests: Ecology and Recovery in a Rapidly Changing World (eds. A. Barton and W. Keeton)
(Invited Book Chapter).
Looney, C.E., A.W. D’Amato, S. Fraver, B.J. Palik, L. Frelich. 2018. Interspecific competition limits
the realized niche of Fraxinus nigra along a waterlogging gradient. Canadian Journal of
Forest Research 48: 1292–1301.
Wei, X., D.J. Hayes, S. Fraver. 2018. Global pyrogenic carbon production during recent decades
has created the potential for a large, long-term sink of atmospheric CO 2. Journal of
Geophysical Research – Biogeosciences 123. DOI: 10.1029/2018JG004490.
2017
Fraver, S., K. Dodds, L.S. Kenefic, R. Morrill, R.S. Seymour, and E. Sypitkowski. 2017. Forest
structure following tornado damage and salvage logging in northern Maine, USA. Canadian
Journal of Forest Research 47:560-564.
Fraver, S., T. Aakala, and A.W. D’Amato. 2017. Dendroecology and coarse woody debris
dynamics. IN: Dendroecology: Tree-ring Analyses Applied to Ecological Studies (eds. A.
Amoroso, L. Daniels, P. Baker, and J.J. Camarero). (Invited Book Chapter).
Ayrey, E., S. Fraver, J.A. Kershaw, A.R. Weiskittel, B. Cook, L.S. Kenefic, B. Roth, and D. Hayes.
2017. Layer Stacking: A novel algorithm for individual tree segmentation from LiDAR point
clouds. Canadian Journal of Remote Sensing 43(1): 16-27.
Curzon, M., A.W. D’Amato, S. Fraver, and B.J. Palik. 2017. Succession, climate and
neighbourhood dynamics influence tree growth over time: an 87-year record of change in a
Pinus resinosa-dominated forest, Minnesota, USA. Journal of Vegetation Science 28: 82-92.
Curzon, M., A.W. D’Amato, S. Fraver, B.J. Palik, A. Bottero, J.R. Foster, and K. Gleason. 2017.
Harvesting influences functional identity and diversity over time in forests of the
northeastern U.S.A. Forest Ecology and Management 400: 93-99.
Carter, D.R., R.S. Seymour, S. Fraver, and A. Weiskittel. 2017. Reserve tree mortality in two
expanding-gap silvicultural systems 20 years after establishment in the Acadian forest of
Maine, USA. Forest Ecology and Management 389:149-157.
Carter, D.R., R.S. Seymour, S. Fraver, and A. Weiskittel. 2017. Effects of multi-aged expanding-
gap silvicultural systems on mature, within-gap reserve tree growth 20 years after
establishment in the Acadian Forest of Maine, USA. Canadian Journal of Forest Research.
47: 1314–1324.
Bottero, A., A.W. D’Amato, B.J. Palik, J.B. Bradford, S. Fraver, M.A. Battaglia, and L.A. Asherin.
2017. Density-dependent vulnerability of forest ecosystems to drought. Journal of Applied
Ecology 54(6): 1605-1614.
Gleason, K.E., J.B. Bradford, A. Bottero, A.W. D’Amato, S. Fraver, B.J. Palik, M.A. Battaglia, L.
Iveson, L. Kenefic, and C. C. Kern. 2017. Competition amplifies drought stress in forests
across broad climatic and compositional gradients. Ecosphere 8(7):e01849.
Kolka, R.K., B.R. Sturtevant, J.R. Miesel, A. Singh, P.T. Wolter, S. Fraver, T.M. DeSutter, and P.A.
Townsend. 2017. Emissions of forest floor and mineral soil carbon, nitrogen and mercury
pools and relationships with fire severity for the Pagami Creek Fire in the boreal forest of
northern Minnesota. International Journal of Wildland Fire 26(4): 296-305.
Costanza K.K.L., W.H. Livingston, D.M. Kashian, R.A. Slesak, J.C. Tardif, J.P. Dech, A.K. Diamond,
J.J. Daigle, D.J. Ranco, J. Neptune, L. Benedict, S. Fraver, M. Reinikainen, and N.W. Siegert.
2017. The precarious state of a cultural keystone species: Tribal and biological assessments
of the future of black ash. Journal of Forestry 115(5): 435-445.
2016
Khakimulina, T., S. Fraver, and I. Drobyshev. 2016. Mixed-severity natural disturbance regime
dominates in an old-growth Norway spruce forest of northwest Russia. Journal of
Vegetation Science 27(2): 400-413.
Puhlick, J.J., S. Fraver, I.J. Fernandez, A.R. Weiskittel, L.S. Kenefic, R.K. Kolka, and M.-C. Gruselle.
2016. Factors influencing organic-horizon carbon pools in mixed-species stands of central
Maine, USA. Forest Ecology and Management 364: 90-100.
Gill, K., A.W. D’Amato, and S. Fraver. 2016. Multiple developmental pathways for range-margin
Pinus banksiana forests. Canadian Journal of Forest Research. 46: 200–214.
Looney, C.E., A.W. D’Amato, S. Fraver, M.R. Reinikainen, and B.J. Palik. 2016. Examining the
influences of tree-to-tree competition and climate on size-growth relationships in hydric,
multi-aged Fraxinus nigra stands. Forest Ecology and Management 375: 238-248.
Puhlick, J.J., A.R. Weiskittel, I.J. Fernandez, S. Fraver, L.S. Kenefic, R.S. Seymour, R.K. Kolka, L.E.
Rustad, and J.C. Brissette. 2016. Long-term influence of alternative forest management
treatments on total ecosystem and wood product carbon storage. Canadian Journal of
Forest Research 46(11): 1404-1412.
Puhlick, J.J., A.R. Weiskittel, S. Fraver, M. Russell, L.S. Kenefic. 2016. Assessing the role of
natural disturbance and forest management on dead wood dynamics in mixed-species
stands of central Maine, USA. Canadian Journal of Forest Research 46: 1092-1102.
2015
Russell, M., S. Fraver, T. Aakala, J. Gove, C.W. Woodall, A.W. D’Amato, and M.J. Ducey. 2015.
Quantifying carbon stores and decomposition in dead wood: A review. Forest Ecology and
Management 350:107-128.
Kuehne, C., A.R. Weiskittel, S. Fraver, and K.J. Puettmann. 2015. Thinning influences structural
heterogeneity, growth, and mortality in Douglas-fir forests. Canadian Journal of Forest
Research 45: 1448-1461.
Forrester, J.A., D.J. Mladenoff, A.W. D’Amato, S. Fraver, D.L. Lindner, N.J. Brazee, and S.T.
Gower. 2015. Temporal trends and sources of variation in carbon flux from coarse woody
debris in experimental forest canopy openings. Oecologia 179: 889–900.
Woodall, C.W., M. Russell, B.F. Walters, A.W. D’Amato, S. Fraver, G. Domke. 2015. Net carbon
flux of dead wood in forests of the Eastern US. Oecologia 177: 861–874.
2014
Fraver, S., A.W. D’Amato, J.B. Bradford, B.G. Jonsson, M. Jönsson, and P.-A. Esseen. 2014. Tree
growth and competition in an old-growth Picea abies forest of boreal Sweden: Influence of
tree spatial patterning. Journal of Vegetation Science 25: 374-385.
Russell, M., C. Woodall, S. Fraver, A.W. D’Amato, G. Domke, and K. Skog. 2014. Residence times
and decay rates for downed woody debris biomass/carbon in eastern US forests.
Ecosystems 17: 765-777.
Svoboda, M., P. Janda, R. Bače, S. Fraver, T. Nagel, and others. 2014. Landscape-level variability
in historical disturbance in primary Picea abies mountain forests of the Eastern Carpathians,
Romania. Journal of Vegetation Science 25: 386-401.
Russell, M., C.W. Woodall, A.W. D’Amato, S. Fraver, and J.B. Bradford. 2014. Linking climate
change and downed woody debris decomposition across forests of the eastern United
States. Biogeosciences 11: 6417-6425.
Brazee, N., D.L. Lindner, A.W. D’Amato, S. Fraver, J.A. Forrester, and D.J. Mladenoff. 2014.
Disturbance and diversity of wood-inhabiting fungi: effects of canopy gaps and downed
woody debris. Biodiversity and Conservation 23: 2155–2172.
Klockow, P.A., A.W. D’Amato, J.B. Bradford, and S. Fraver. 2014. Nutrient concentrations in
coarse and fine woody debris in Populus tremuloides-dominated forests, northern
Minnesota, USA. Silva Fennica 48(1): Article ID 962.
Reinikainen, M., A.W. D’Amato, J.B. Bradford, and S. Fraver. 2014. Influence of stocking, site
quality, stand age, low-severity canopy disturbance, and forest composition on sub-boreal
aspen mixedwood carbon stocks. Canadian Journal of Forest Resources 44: 230-242.
Kolka, R., B. Sturtevant, P. Townsend, J. Miesel, P. Wolter, S. Fraver, and T. DeSutter. 2014.
Forest floor and upper mineral soil carbon, nitrogen and mercury pools shortly after fire and
comparisons utilizing fire severity indices. Soil Science Society of America Journal 78(S1):
S58-S65.
2013
Fraver, S., A.M. Milo, J.B. Bradford, A.W. D’Amato, L. Kenefic, B.J. Palik, C.W. Woodall, and J.
Brissette. 2013. Woody debris volume depletion through decay: Implications for biomass
and carbon accounting. Ecosystems 16(7): 1262-1272.
Lindbladh, M., S. Fraver, J. Edvardsson, and A. Felton. 2013. Past forest composition, structures
and processes – How paleoecology can contribute to forest conservation. Biological
Conservation 168: 116-127.
Aakala T., S. Fraver, A.W. D’Amato, and B.J. Palik. 2013. Influence of competition and age on
tree growth in structurally complex old-growth forests in northern Minnesota, USA. Forest
Ecology and Management 308: 128-135.
Silver, E.J., S. Fraver, A.W. D’Amato, T. Aakala, and B.J. Palik. 2013. Long-term mortality rates
and spatial patterns in an old-growth Pinus resinosa forest. Canadian Journal of Forest
Research 43: 809–816.
D’Amato, A.W. J.B. Bradford, S. Fraver, and B.J. Palik. 2013. Effects of thinning on drought
vulnerability and climate response in north temperate forest ecosystems. Ecological
Applications 23(8): 1735-1742.
Russell, M., C. Woodall, S. Fraver, and A.W. D’Amato. 2013. Estimates of coarse woody debris
decay class transitions for forests across the eastern United States. Ecological Modelling 251: 22-31.
Berger, A.L., B. Palik, A.W. D’Amato, S. Fraver, J.B. Bradford, K. Nislow, D. King, and R.T. Brooks.
2013. Ecological impacts of energy-wood harvests: Lessons from whole-tree harvesting and
natural disturbance. Journal of Forestry 111(2): 139-153.
Silver, E.J., A.W. D’Amato, S. Fraver, B.J. Palik, and J.B. Bradford. 2013. Structure and
development of old-growth, unmanaged second-growth, and extended rotation Pinus
resinosa forests in Minnesota, USA. Forest Ecology and Management 291: 110-118.
2012
Fraver, S. and B.J. Palik. 2012. Stand and cohort structures of old-growth Pinus resinosa-
dominated forests of northern Minnesota, USA. Journal of Vegetation Science 23: 249-259.
Aakala, T., S. Fraver, B.J. Palik, and A.W. D’Amato. 2012. Spatially random mortality in old-
growth red pine forests of northern Minnesota. Canadian Journal of Forest Research 42:
899-907.
Bradford, J.B., S. Fraver, A.M. Milo, A.W. D’Amato, B.J. Palik, and D.J. Shinneman. 2012. Effects
of multiple interacting disturbances and salvage logging on forest carbon stocks. Forest
Ecology and Management 267: 209-214.
Mitchell, C.P.J., R.K. Kolka, and S. Fraver. 2012. Singular and combined effects of blowdown,
salvage logging, and wildfire on forest floor and soil mercury pools. Environmental Science &
Technology 46: 7963-7960.
Brazee, N., D.L. Lindner, S. Fraver, A.W. D’Amato, and A.M. Milo. 2012. Wood-inhabiting,
polyporoid fungi in aspen-dominated forests managed for biomass in the U.S. Lake States.
Fungal Ecology 5: 600-609.
Powers, M.D., R.K. Kolka, J.B. Bradford, B.J. Palik, S. Fraver, and M.F. Jurgensen. 2012. Carbon
storage across a chronosequence of thinned and unmanaged red pine stands. Ecological
Applications 22(4): 1297-1307.
Reinikainen, M., A.W. D’Amato, and S. Fraver. 2012. Repeated insect outbreaks promote multi-
cohort aspen mixedwood forests in the Lake States, USA. Forest Ecology and Management
266: 148-159.
Svoboda, M., P. Janda, T. Nagel, S. Fraver, J. Rejzek, and R. Bače. 2012. Disturbance history of
an old-growth sub-alpine Picea abies stand in the Bohemian Forest, Czech Republic. Journal
of Vegetation Science 23: 86-97.
2011
Fraver, S., T. Jain, J.B. Bradford, A.W. D’Amato, D. Kastendick, B.J. Palik, and D. Shinneman.
2011. The efficacy of salvage logging in reducing subsequent fire severity in conifer-
dominated forests of Minnesota, USA. Ecological Applications 21(6): 1895-1901.
Fraver, S., J.B. Bradford, and B.J. Palik. 2011. Improving tree age estimates derived from
increment cores: a case study of Pinus resinosa. Forest Science 57:164-170.
Jönsson, M., S. Fraver, and B.G. Jonsson. 2011. Spatio-temporal variation of coarse woody
debris input in woodland key habitats in central Sweden. Silva Fennica 45(5): 957-967.
D’Amato, A.W., S. Fraver, B. Palik, J. B. Bradford, and L. Patty. 2011. Singular and interactive
effects of blowdown, salvage logging, and wildfire in sub-boreal pine systems. Forest
Ecology and Management 262: 2070-2078.
Saunders, M.R., S. Fraver, and R.G. Wagner. 2011. Nutrient concentration of down woody
debris in mixedwood forests in central Maine, USA. Silva Fennica 45(2): 197-210.
Hanberry, B., S. Fraver, J. Yang, H.S. He, and B.J. Palik. 2011. Spatial pattern corrections and
sample sizes for forest density estimates of historical tree surveys. Landscape Ecology 26:
59-68.
D’Amato, A.W., J. Bradford, S. Fraver, and B.J. Palik. 2011. Forest management for mitigation
and adaptation: insights from long-term silvicultural experiments. Forest Ecology and
Management 262: 803-816.
2010 and prior
Svoboda, M., S. Fraver, P. Janda, R. Bače, and J. Zenáhlíková. 2010. Natural development and
regeneration of a Central European montane spruce forest. Forest Ecology and
Management 260: 707-714.
Bradford, J.B., A.W. D’Amato, B.J. Palik, and S. Fraver. 2010. A new method for evaluating forest
thinning: growth dominance in managed Pinus resinosa stands. Canadian Journal of Forest
Research 40: 843-849.
Powers, M.D., B.J. Palik, J.B. Bradford, S. Fraver, C.R. Webster. 2010. Thinning method and
intensity influence long-term mortality trends in a red pine forest. Forest Ecology and
Management 260: 1138-1148.
Fraver, S., A.S. White, and R.S. Seymour. 2009. Natural disturbance in an old-growth landscape
in northern Maine, USA. Journal of Ecology 97: 289-298.
Jönsson, M., S. Fraver, and B.G. Jonsson. 2009. Forest history and the development of old-
growth characteristics in Swedish spruce forests. Journal of Vegetation Science 20: 91-106.
Fraver, S., B.G. Jonsson, M. Jönsson, and P.-A. Esseen. 2008. Demographics and disturbance
history of a boreal old-growth Picea abies forest. Journal of Vegetation Science 19: 789-798.
Fraver, S., A. Ringvall, and B.G. Jonsson. 2007. Refining volume estimates of down woody
debris. Canadian Journal of Forest Research 37: 627-633.
Fraver, S., R.S. Seymour, J. H. Speer, and A.S. White. 2007. Dendrochronological reconstruction
of spruce budworm outbreaks in northern Maine. Canadian Journal of Forest Research 37:
523-529.
Jönsson, M., S. Fraver, B.G. Jonsson, M. Dynesius, M. Rydgård, and P.-A. Esseen. 2007. Eighteen
years of tree mortality and structural change in an experimentally fragmented Norway
spruce forest. Forest Ecology and Management 242: 306-313.
Fraver, S. and A.S. White. 2005. Disturbance dynamics of old-growth Picea rubens forests of
northern Maine. Journal of Vegetation Science 16: 597-610.
Fraver, S. and A.S. White. 2005. Identifying growth releases in dendrochronological studies of
forest disturbance. Canadian Journal of Forest Research 35(7): 1648-1656.
Kenefic, L.S., A.S. White, A.R. Cutko, and S. Fraver. 2005. Reference stands for silvicultural
research: a Maine perspective. Journal of Forestry 103(5): 363-367.
Brokaw, N.V.L., S. Fraver, J.S. Grear, J. Thompson, J.K. Zimmerman, R.B. Waide, E.M. Everham
III, S.P. Hubbell, and R.B. Foster. 2004. Disturbance and canopy structure in two tropical
forests. Pages 177-194 in E.C. Losos and E.G. Leigh, Jr., editors. Tropical Forest Diversity and
Dynamism: Findings from a Large-Scale Plot Network. University of Chicago Press.
Lara, A., A. Wolodarsky-Franke, J.C. Aravena, M. Cortés, S. Fraver, and F. Silla. 2003. Fire regimes
and forest dynamics in the Lake Region of South-Central Chile. Pages 323-342 in T.T. Veblen,
W.L. Baker, G. Montenegro, and T.W. Swetnam, editors. Fire and Climate Change in Temperate
Ecosystems of the Western Americas. Springer-Verlag New York, Inc.
Fraver, S., R.G. Wagner, and M. Day. 2002. Dynamics of coarse woody debris following gap
harvesting in the Acadian forest of central Maine, USA. Canadian Journal of Forest Research
32(12): 2094-2105.
Silla, F., S. Fraver, A. Lara, T.R. Allnutt, and A. Newton. 2002. Regeneration and stand dynamics
of Fitzroya cupressoides (Cupressaceae) forests of southern Chile’s Central Depression.
Forest Ecology and Management 165(1): 213-224.
Lara, A., S. Fraver, J.C. Aravena, and A. Wolodarsky-Franke. 1999. Fire and the dynamics of Fitzroya
cupressoides (alerce) forests of Chile’s Cordillera Pelada. EcoScience 6(1): 100-109.
Fraver, S., M.E. González, F. Silla, A. Lara, and M. Gardner. 1999. Composition and structure of
remnant Fitzroya cupressoides forests of Southern Chile’s Central Depression. Journal of the
Torrey Botanical Society 126(1): 49-57.
Fraver, S., N.V.L. Brokaw, and A.P. Smith. 1998. Delimiting the gap phase in the growth cycle of
a Panamanian forest. Journal of Tropical Ecology 14(5): 673-681.
González, M.E., C. Donoso, and S. Fraver. 1997. Results from mixed plantations of Eucryphia
cordifolia, Laurelia sempervirens and Aextoxicon punctatum established in areas disturbed
by mass flowering of the bamboo, Chusquea quila. Bosque 18(1): 53-59.
Shear, T.H., T.J. Lent, and S. Fraver. 1996. Comparison of restored and mature bottomland
hardwood forests of southwestern Kentucky. Restoration Ecology 4(2): 111-123.
Fraver, S. 1994. Vegetation responses along edge-to-interior gradients in the mixed hardwood
forests of the Roanoke River Basin, North Carolina. Conservation Biology 8(3): 822-832.
Fraver, S. 1992. The insulating value of serotinous cones in protecting pitch pine (Pinus rigida)
seeds from high temperatures. Journal of the Pennsylvania Academy of Science 65(3): 112-
116.